The table below presents genomic information on specimens in all the genera EXCEPT Triticum, Aegilops, and Amblyopryum which is on another page. The symbols used follow the recommendations of Wang et al. (1994). Notice that these authors recommended using E, not J, for the genome in T. elongatum and its allies (Thinopyrum as used on this web site), in order be consistent with previous use by wheat breeders. For information on this page I am indebted to Richard Wang and Bjorn Salomon. The information on the Triticum-Aegilops-Amblyopyrum page came from Richard Wang and Michael van Slageren.
The table below shows the genomic constitution of individual taxa sorted by genomes within affiliated groups of genera. For further information about these records, click here. To move to groups, use the following links: St genome species, E genome species, N genome species. Because of the way in which the species are grouped, it is impossible to make this as easy to use as might be desirable. One day I shall split it into multiple tables, each table including all taxa known to have a particular genome. When this happens, species polygenomic species will appear in more than one table.
The last column contains the generic name that should be used if adopting the principle of genomic classification. It is based on the recommendations of Yen et al. . 2005, with one modification - the use of Thinopyrum rather than Lophopyrum when the two genera are combined in order to conform to the requiremens of the International Code of Botanical Nomenclature. Among the the principles Yen et al. followed in making their recommendations was that the treatment of genera should reflect both phylogenetic relationships and convenience and that force of habit should be considered, whenever possible, to avoid chaos. They acknowledge that their system is incomplete and does not resolve all problems. Prof. Yen sent me (Nov. 17, 2005) a working synopsis of Roegneria, for which I thank him. That information is being incorporated into the page for Roegneria. This page is designed to show which taxa have been genomically characterized and where the information was published.
Taxon |
Genomes |
Reference |
Genomic genus, see Yen et al. . 2005 |
| Agropyron | P | Agropyron | |
| Crithopsis | K | ||
| Dasypyrum breviaristatum | VbVb | Galasso et al. 1997; Ohta and Morishita 2001 | |
| Dasypyrum villosum | V | Galasso et al. 1997 | |
| Pseudoroegneria | St | Pseudoroegneria | |
| Pseudoroegnerai strigosa subsp. aegilopoides | St | Yu et al. (2008) | |
| Pseudoroegneria gracillima | St | Yu et al. (2008) | |
| Pseudoroegneria stipifolia | St | Yu et al. (2008) | |
| Pseudoroegnera strigosa subsp. strigosa | St | Yu et al. (2008) | |
| Pseudoroegneria libanotica | St | ||
| Pseudoroegneria tauri = Douglasdewey wangii | StP | Wang et al. (1986) | Douglasdeweya |
| Roegneria magnicaespes | St | Ding et al. 2004; Ding et al. 2005; Zhang et al. 1999 | ;Pseudoroegneria |
| Pseudoroegneria alashanica | St | Zhang et al. 1999 ; Zhang et al. 2002; Ding et al. 2004 | Pseudoroegneria |
| Roegneria elytrigioides | St | Lu 1994; Ding et al. 2004 | Pseudoroegneria |
| Pseudoroegneria cognata | St | Dewey 1981 | Pseudoroegneria |
| Elymus agropyroides | StH | Dewey 1970b. | Elymus |
| Elymus alaskanus | StH | Oergaard & Anamthawat-Jonsson 2001 | Elymus |
| Elymus albicans | StH | Dewey 1970a. | Elymus |
| Elymus canadensis | StH | Dewey 1970; Mason-Gamer 2001 | Elymus |
Elymus caninus Type of Elymus sect. Goulardia [sect. name from 1941; Cynapoa is an older sectional name] |
StH | Dewey 1970; Redinbaugh et al. 2000 | Elymus |
| StH | Dewey 1974; Redinbaugh et al. 2000; Oergaard & Anamthawat-Jonsson 2001. | ||
| Elymus dasystachys | StH | Dewey 1967 [Need to check this] | Elymus |
| StH | Dewey 1970; Redinbaugh et al. 2000; | ||
| Elymus elymoides | StH | Dewey 1964; Redinbaugh et al. 2000; Mason-Gamer 2001; Larson et al. 2003. | |
| Elymus hystrix | StH | Mason-Gamer 2001; Mason-Gamer et al. 2002. | Elymus |
| Elymus glaucus | StH | Dewey 1965; Mason-Gamer 2001; Mason-Gamer et al. 2002. | Elymus |
| Mason-Gamer et al. 2002. | |||
| Elymus lanceolatus | StH | Dewey 1967 (as Agropyron dasystachyum); Mason-Gamer 2001; Mason-Gamer et al. 2002. | Elymus |
| Elymus magellanicus | StH | Jensen 1993. | Elymus |
| Elymus multisetus | StH | Larson et al. 2003.. | Elymus |
| Elymus mutabilis | StH | Dewey 1979; Redinbaugh et al. 2000. | Elymus |
| Elymus pringlei | StH | Dewey 1976. | Elymus |
| Elymus scribneri | StH | Dewey 1967. | Elymus |
| Elymus sibiricus | StH | Dewey 1974. | Elymus |
| Elymus subsecundus | StH | Dewey 1966. | Elymus |
| Elymus tilcarensis | StH | Dewey 1977. | Elymus |
| Elymus trachycaulus | StH | Dewey 1977; Redinbaugh et al. 2000 | Elymus |
| StH | Mason-Gamer et al. 2002. | ||
| Elymus virginicus | StH | Mason-Gamer 2001; Mason-Gamer et al. 2002. | Elymus |
| Elymus wawawaiensis | StH | Mason-Gamer 2001; Mason-Gamer et al. 2002. | Elymus |
| Elymus hoffmannii | StStH | Jensen & Asay 1996. | |
| Elymus transhyrcanus | StStH | Dewey 1972; Redinbaugh et al. 2000. | |
| Elymus repens | StStH | Assadi et Runemark 1994; Vershinin et al.1994 | |
| StStH | Assadi et Runemark 1994; Vershinin et al.1994 | ||
| StStH | Redinbaugh et al. 2000 | ||
| XXH | Oergaard & Anamthawat-Jonsson 2001 | ||
| StH(V-E-Ta-X) | Mason-Gamer et al. 2002. | ||
| Elymus patagonicus | StHH | Redinbaugh et al. 2000. | |
Elymus dahuricus Type of Elymus sect. Turczaninovia [sect. epithet 1932] |
StHY | Redinbaugh et al. 2000. | Campeiostachys |
| Elymus drobovii | StHY | Dewey 1980 | Campeiostachys |
| Elymus himalayanus | StHY | Lu and von Bothmer 1992. | Campeiostachys |
| Elymus kamoji | StHY | Zhang et al. 1999. | Campeiostachys |
| Elymus nutans | StHY | Lu 1993. | Campeiostachys |
| Elymus schrenkianus | StHY | Lu and von Bothmer 1992. | Campeiostachys |
| Elymus tsukushiensis | StHY | Lu et von Bothmer 1990a; Redinbaugh et al. 2000. | Campeiostachys |
| Elymus caesifolius | StStY | Svitashev et al. 1998 | |
| Elymus altissimus | StY | Lu and Salomon1992; Lu and von Bothmer 1993a. | Roegneria |
| Elymus anthosachnoides | StY | Lu and Salomon1992; Lu et von Bothmer 1993a. | Roegneria |
| Elymus barbicallus | StY | Lu and Salomon1992. | Roegneria |
| Elymus breviglumis | StY | Lu and Salomon1992. | Roegneria |
| Elymus brevipes | StY | Lu et von Bothmer 1993a. | Roegneria |
| Elymus cacuminus | StY | Lu and Salomon 1993b. | Roegneria |
Elymus caucasicus Type of Elymus sect. Clinelymopsis (1934) |
StY | Jensen and Wang 1991; Lu and von Bothmer 1993b; Redinbaugh et al. 2000. | Roegneria |
| Elymus caucasicus | StY | Jensen and Wang 1991; Lu et von Bothmer 1993b; Redinbaugh et al. 2000 | Roegneria |
| Elymus ciliaris | StY | Zhou et al. 1999; Redinbaugh et al. 2000; Mason-Gamer et al. 2002. | Roegneria |
| Elymus ciliaris | StY | Zhou et al. 1999; Redinbaugh et al. 2000 | Roegneria |
| Elymus dolichaterum | StY | Lu et Salomon1992. | Roegneria |
| Elymus fedtschenkoi | StY | Liu et Dewey 1983. | Roegneria |
| Elymus gmelinii | StY | Jensen et Hatch 1989; Lu et Salomon1992. | Roegneria |
| Elymus grandis | StY | Zhang et al. 1998a; Svitashev et al. 1998. | Roegneria |
| Elymus jacquemontii | StY | Lu 1993. | Roegneria |
| Elymus longearistatus | StY | Jensen et Wang 1991; Lu et von Bothmer 1993b. | Roegneria |
| Elymus nevskii | StY | Lu et Salomon1992; Redinbaugh et al. 2000 | Roegneria |
| Elymus nipponicus | StY | Lu et Salomon1992. | Roegneria |
| Elymus panormitanus | StY | Lu et Salomon1992; Redinbaugh et al. 2000 | Roegneria |
| Elymus parviglumis | StY | Lu et von Bothmer 1990b; Lu et Salomon 1993a. | Roegneria |
| Elymus pendulinus | StY | Jensen 1990a; Zhou et al. 1999; Redinbaugh et al. 2000 | Roegneria |
| Elymus praeruptus | StY | Salomon 1993. | Roegneria |
| Elymus pseudonutans | StY | Lu et von Bothmer 1990b; Lu et Salomon1992. | Roegneria |
| Elymus retroflexus | StY | Lu et Salomon 1993b. | Roegneria |
| Elymus shandongensis | StY | Lu et Salomon1992. | Roegneria |
| Elymus sclerus | StY | Lu et Salomon1992. | Roegneria |
| Elymus semicostatus | StY | Lu et Salomon 1991, 1993a; Redinbaugh et al. 2000; | Roegneria |
| Elymus strictus | StY | Lu et Salomon1992; Redinbaugh et al. 2000 | Roegneria |
| Elymus tenuispicus | StY | Zhou et al. 1999 | Roegneria |
| Elymus tibeticus | StY | Lu et Salomon1992, 1993a. | Roegneria |
| Elymus validus | StY | Lu et Salomon1992. | Roegneria |
| Elymus varius | StY | Lu et Salomon1992. | Roegneria |
| Elymus yangiae | StY | Lu et Salomon1992; Lu et von Bothmer 1993a. | Roegneria |
| Elymus borianus | StStY | Svitashev et al. 1998. | |
| Elymus caesifolius | StStY | Svitashev et al. 1998. | |
| Elymus glaucissimus | StStY | Lu et al. 1993; Redinbaugh et al. 2000 | |
| Elymus tschimganicus | StStY | Lu et al. 1993; Jensen et al. 1994; Redinbaugh et al. 2000 | |
| Kengyilia | StPY | Yen et Yang 1990 | Kengyilia |
| Elymus alatavicus | StPY | Jensen et al. 1986; Redinbaugh et al. 2000 | Kengyilia |
| Elymus batalinii | StPY | Jensen et al. 1986; Redinbaugh et al. 2000 | Kengyilia |
| Elymus gobicola | StPY | Zhang et al. 2000. | Kengyilia |
| Elymus grandiglumis | StPY | Jensen 1990b; Redinbaugh et al. 2000 | Kengyilia |
| Elymus hirsuta | StPY | Zhang et al. 1999. | Kengyilia |
| Elymus kengii | StPY | Jensen 1990b; Redinbaugh et al. 2000 | Kengyilia |
| Elymus kokonoricus | StPY | Yang et al. 1992; Jensen 1996. | Kengyilia |
| Elymus laxiflora | StPY | Zhang et al. 1998b. | Kengyilia |
| Elymus melantherus | StPY | Yang et al. 1992; Jensen 1996. | Kengyilia |
| Elymus muticus | StPY | Zhang et al. 1999. | Kengyilia |
| Elymus retusus | StPY | Yang et al. 1992. | Kengyilia |
| Elymus rigidulus | StPY | Yang et al. 1992; Zhang et al. 1999. | Kengyilia |
| Elymus stenachyrus | StPY | Yang et al. 1992. | Kengyilia |
| Elymus thoroldianus | StPY | Yang et al. 1992; Jensen 1996. | Kengyilia |
| Elymus zhaosuensis | StPY | Zhang et al. 2000. | Kengyilia |
| Elymus apricus | StYW | Stewart et al. 2005. | Anthosachne |
| Douglasdeweya deweyi (Voucher SAUTI | StP | Jensen et al. 1992; Yen et al. 2005. | Douglasdeweya |
| Douglasdeweya wangii = Pseudoroegneria tauri (Voucher SAUTI 9109; grown from PI 401328) |
StP | Wang et al. . 1986; Yen et al. 2005. | Douglasdeweya |
Pseudoroegneria pertenuis |
StP | Wang et al. . 1986; Assadi 1995 but see Yen et al. 2005 | |
| Pascopyrum smithii | StHNsXm | Zhang and Dvorak 1991 | |
| Elytrigia caespitosa subsp. caespitosa | EeSt | Yu et al. 2008 | Thinopyrum |
| Pseudoroegneria geniculata subsp. scythica | EeSt | Liu et Wang 1992 | Thinopyrum |
| Pseudoroegneria tauri (Voucher PI 343188) |
St | Dewey 1974 | |
| Elytrigia caespitosa subsp. caespitosa [combination in Thinopyrum is invalid] | EeSt | Liu et Wang 1989, 1993b | Thinopyrum |
| Elytrigia caespitosa subsp. nodosa [combination not made] | EeSt | Liu et Wang 1993b | Thinopyrum |
| Thinopyrum intermedium | EbEeSt | Liu et Wang 1993b; Xu et Conner 1994 | Thinopyrum |
| EEsSt | Chen et al. 1998 | ||
| EeSt(V-Eb-R) | Kishii et al. 2005. | ||
| Thinopyrum ponticum | EEEStSt | Zhang et al. 1996. | Thinopyrum |
| EEEEE | Muramatsu 1990; Wang et al. 1991. | ||
| EEEEsEs | Chen et al. 1998. | ||
| Psammopyrum athericum | EStP or ELP | Ellneskog-Staam et al. 2001; .Ellneskog-Staam et al. 2003. | Thinopyrum [combination not made] |
| Psammopyrum pungens | EStStP or EStLP | Ellneskog-Staam et al. 2003. | Thinopyrum |
| Thinopyrum elongatum | Ee | Wang 1985 | Thinopyrum |
| Thinopyrum bessarabicum | Eb | Wang 1985 | Thinopyrum |
| Thinopyrum curvifolium | EbEb | Liu et Wang 1993a | Thinopyrum |
| EcEcu | McGuire 1984. | ||
| Thinopyrum distichum | EbEe | Liu et Wang 1993a | Thinopyrum |
| Ed1Ed2 | Pienaar et al. 1988; Liu et Wang 1993a. | ||
| Thinopyrum junceiforme | EbEe | Liu et Wang 1992; Refoufi et al. 2001 | Thinopyrum |
| Thinopyrum junceum | EbEbEe | Liu et Wang 1993a | Thinopyrum |
| Thinopyrum sartorii | EbEe | Liu et Wang 1992 | Thinopyrum |
| Thinopyrum scirpeum | EeEe | Liu et Wang 1993a | Thinopyrum |
| EsEsc | McGuire 1984. | ||
| Thinopyrum pycnanthum = Elytrigia pycnantha | EStP | Refoufi et al. 2001 | Thinopyrum |
| Crithopsis | K | ||
| Elymus rectisetus | StYW | Torabinejad et Mueller 1993; Redinbaugh et al. 2000 | Anthosachne (in litt. 2005/10/30) |
| Elymus scaber | StYW | Torabinejad et Mueller 1993; Redinbaugh et al. 2000 | Anthosachne (in litt. 2005/10/30) |
| Australopyrum | W | Australopyrum | |
| Elymus scabrifolius | ?H | Linde-Laursen et Seberg 2002 | |
| Eremopyrum | FXe | Sakamoto 1979; Frederiksen and Bothmer 1995 | |
| Festucopsis | L | ||
| Henrardia | O | ||
| Heteranthelium | Q | ||
| Hordeum bulbosum | I | ||
| Hordeum marinum | Xa | Bothmer et al. . 1986 | |
| Hordeum murinum | Xu | Bothmer et al. . 1987, 1988a, 1998b | |
| Hordeum vulgare | I | ||
| Hordeum, other diploid species | H | ||
| Psathyrostachys | Ns | ||
| Hordelymus europaeus | Ns? | Svitashev et al. 1998. | |
| XoXr | Bothmer et al. . 1994 | ||
| Ns(ancestor of St and L genomes) | Petersen, G. and O. Seberg. 2008 | ||
| Leymus aemulis | NsXm | Sun et al. . 1994. | |
| Leymus ambiguus | NsXm | Dewey 1976 | |
| Leymus arenarius | NsNsXmXm | Wang et Jensen 1994. | |
| Leymus cinereus | NsXm | Wu et al. 2003. | |
| Leymus coreanus | NsXm | Jensen et Wang 1997. | |
| Elymus duthiei | NsXm | Svitashev et al. 1998 | |
| Leymus erianthus | NsXm | Dubcovsky et al. 1997. | |
| Leymus flavescens | NsXm | Hole et al. 1999 | |
| Elymus komarovii | NsXm | Svitashev et al. 1998 | |
| Leymus mendocinus | NsXm | Dubcovsky et al. 1997. | |
| Leymus mollis | NsXm | Wang and Hsiao 1984. | |
| Leymus triticoides | NsXm | Wu et al. 2003. | |
| Leymus ugamicus | NsXm | Dewey 1980b. | |
| Leymus | NsXm | Zhang et Dvorak 1991; Wang et Jensen 1994 | |
| Leymus californicus | Ns? | Jensen et Wang 1997; Dubcovsky et al. . 1997; | |
| Ns? | Svitashev et al. 1998; Mason-Gamer 2001. | ||
| NsXm | Jensen et Wang 1997. | ||
| Pascopyrum smithii | StHNsXm | Zhang and Dvorak 1991 | |
| Peridictyon sanctus | Xp | Seberg et al. . 1991 | |
| Australopyrum | W | Assigned by Löve 1984 | |
| Elymus enysii | HW | Stewart et al. 2005 | Stenostachys |
| Secale | R | ||
| Taeniatherum | Ta | ||
| Pseudoroegneria kosaninii | not known |
Pseudoroegneria pertenuis: Assadi (1994, 1995) identified plants from Azerbaijan and those grown from the US accessions PI 40328 and 401330 as Elymus pertenuis = Pseudoroegneria pertenuis. Yen et al. . (2005) concluded that plants grown from seed of the two accessions were not the same as the type of pertenuis but belonged to a previously undescribed species, which they placed in their new genus Douglasdeweya as D. wangii. The accessions were originally identified as belonging to Pseudoreogneria tauri. According to Yen et al. . (2005), the genomic constitution of the pertenuis taxon is unknown.