| Jacques Cayouette Stephen J. Darbyshire |
Plants perennial; rhizomatous, sometimes producing
aquatic leaves when submerged. Culms (5)10-100 cm, erect, glabrous,
rooting at the lower nodes; nodes glabrous. Sheaths of
aquatic leaves closed to near the apices, translucent, pale pinkish-brown; collars inconspicuous; ligules acute; blades pinkish-brown. Sheaths
of aerial leaves usually closed for over 1/2 of their length, sometimes
open to the base, opaque, green to olive-green, glabrous; collars conspicuous
as a zone of contrasting color; auricles absent; ligules membranous,
truncate, lacerate; blades usually flat, glabrous, upper blades
conspicuously longer than the lower blades. Inflorescences open
panicles; branches stiff and ascending to pendulous, glabrous. Spikelets pedicellate,
somewhat laterally compressed; florets (1)2-7(9); rachillas prolonged
beyond the uppermost floret, glabrous; disarticulation above the
glumes. Glumes subequal, broadly lanceolate to ovate, membranous
to subcoriaceous, glabrous, 1-3(-5)-veined, acute to obtuse; calluses short,
blunt, bearded to almost glabrous; lemmas ovate, glabrous, membranous
to subcoriaceous, with 3(5) obscure veins, apices entire, obtuse; paleas subequal
to the lemmas; lodicules 2, free, glabrous, toothed or entire; anthers 3; ovaries glabrous. Caryopses falling
free; hila broadly ovate, 1/6-1/5 the length of the caryopses. x =
7. From the Greek arktos, north, and philia, loving.
Arctophila is a monotypic, but highly polymorphic, holarctic genus closely related to Dupontia.
1. Arctophila fulva (Trin.) Andersson
Pendant Grass, Arctophile Fauve
Culms (5)10-80(100) cm. Ligules (1)2-6(8) mm; blades 2-23
cm long, 1-5(10) mm wide. Panicles 3-20 cm long, (1.5)3-11 cm wide. Spikelets 2.5-7(8)
mm; florets (1)2-7(9). Glumes 1.5-4(5) mm; lower glumes exceeded
by the lowest floret; upper glumes shorter to longer than the lowest
floret; lemmas 2.5-4 mm; paleas (1)1.8-4 mm; anthers 1.2-3
mm. Caryopses 1.5-2.2 mm. 2n = 42, 63.
Arctophila fulva grows as an emergent species in shallow, standing
water or along slow-moving streams, wet meadows, marshes, and saturated
soils of low arctic and subarctic regions, where it often forms pure stands.
It is one of the few grasses that develop aquatic leaves. Field observations
indicate that under some environmental conditions, A. fulva can
propagate vegetatively from detached stems that have over-wintered (Aiken
and Buck 2002). In the Flora region, it grows from Alaska through
the Yukon, Northwest Territories, Nunavut, Ontario, Quebec, and Labrador
to Greenland. Its range extends across Eurasia to arctic Scandinavia. It
forms a sterile hybrid, ×Arctodupontia scleroclada,
with Dupontia fisheri.