| John R. Reeder |
Plants annual; viscid, aromatic. Culms
with solid, pithy interiors, often with 5 or more nodes. Leaves with little
or no distinction between sheath and blade; ligules absent; blades
of upper cauline leaves similar in length to those below; siliceous cells of
the leaf epidermis absent or irregular to dumbbell-shaped; microhairs of
the blades bicellular, small, sunken, "mushroom-button" shaped. Inflorescences
terminal, narrow, cylindrical to clavate or capitate, usually dense panicles or
spikes, spikelets sessile or shortly pedicellate; rachillas tardily disarticulating
between the florets. Spikelets with 4-25(40) florets. Glumes entire,
denticulate, toothed, or absent; lemmas with 5-17 conspicuous veins; paleas
subequal to or slightly shorter than the lemmas, well-developed, keels glabrous;
lodicules absent or 2, obscure, rounded, truncate to slightly emarginate;
anthers 3. Caryopses laterally compressed; hila large, punctate,
basal; embryos 3/4 or more as long as the caryopses. x = 10.
The tribe Orcuttieae includes only three genera and nine species, all of
which are endemic to vernal pools and similar habitats in California and Baja
California, Mexico. Its member employ C4 photosynthesis in al their leaves, judging by the carbon isotype ratios, but the aquatic leaves of most species of Orcuttia do not exhibit Kranz leaf anatomy whereas those of Tuctoria and Neostapfia do, as do the floating and aerial leaves (Boykin et al. 2008).
1 |
Lemmas deeply
cleft into 5 mucronate or awn-tipped teeth, the teeth 1/3 as long as to
equaling the lemma bodies; spikelets distichously arranged ..... 19.01
Orcuttia |
Lemmas entire or denticulate, often with
a central mucro; spikelets spirally arranged (2) |
|
Inflorescences clavate,
often partially enclosed at maturity; spikelets laterally compressed,
with glumes; lemmas rectangular, not translucent between the veins, the
apices mucronate, otherwise entire or denticulate; caryopses without a
viscid exudate, the embryo visible through the pericarp ..... 19.02 Tuctoria |
|
Inflorescences cylindrical, usually completely
exposed at maturity; spikelets dorsally compressed, without glumes; lemmas
flabellate, translucent between the veins, the apices ciliolate; caryopses
covered with a viscid exudate, the embryos obscured by the pericarp .....
19.03 Neostapfia |