| Claus Baden† |
Plants perennial; cespitose, forming dense to
loose clumps, sometimes stoloniferous, sometimes rhizomatous. Culms 15-120
cm, erect or decumbent. Basal sheaths closed, becoming somewhat
fibrillose; ligules 0.2-0.3 mm, membranous; auricles sometimes
present. Inflorescences spikes; disarticulation at the
rachis nodes above the spikelets. Spikelets sessile, 2-3 per
node, with 1-2(3) florets, often with additional reduced florets distally. Glumes (including
awns) equal to unequal, (3.5)4.2-48.5(65) mm, subulate, scabrous to
pubescent, obscurely 1-veined, not united; lemmas 5.5-14.3 mm,
narrowly elliptic, rounded, glabrous or pubescent, 5-7-veined, veins
often prominent apically, apices sharply acute to awned, sometimes
with a minute tooth on either side of the awn base, awns (0.9)1.4-34
mm, ascending to slightly divergent, sometimes with a violet tinge; paleas equaling
or slightly longer than the lemmas, membranous, scabrous to pilose
on and sometimes between the keels, bifid; anthers 3, 3-6.8(7)
mm, yellow or violet; lodicules 2, acute, entire, ciliate. Caryopses pubescent
apically, tightly enclosed by the lemmas and paleas at maturity. x =
7. Name from the Greek psathyros, fragile, and stachys,
spike.
Psathrostachys has eight species, all of which are native to arid
regions of central Asia, from eastern Turkey to Xinjiang Province, China
and eastern Siberia. One species has been introduced into North America
as a potential forage species.
Psathyrostachys is very similar to Leymus,
particularly the cespitose species of Leymus. Both genera are monogenomic,
having only the N genome.
Bödvarsdóttir and Anamthawat-Jónsson (2003) could not
find any molecular probes that would distinguish between the two genera.
1. Psathyrostachys juncea (Fisch.) Nevski
Russian Wildrye
Plants densely cespitose. Culms (20)30-80(120) cm, erect
or decumbent at the base, mostly glabrous, pubescent below the spikes. Basal
sheaths glabrous, grayish-brown, old sheaths more or less persistent; blades (1)2.5-18(30)
cm long, (1)5-20 mm wide, flat or involute, abaxial surfaces smooth or
scaberulous, often glaucous; auricles 0.2-1.5 mm; ligules 0.2-0.3
mm. Spikes (3)6-11(16) cm long, 5-17 mm wide, erect; rachises hirsute
on the margins, puberulent elsewhere; internodes 3.5-6 mm. Spikelets 7-10(12)
mm (excluding the awns), strongly overlapping, the lateral spikelets slightly
larger than the central spikelets. Glumes (3.5)4.2-9.4 mm, subulate,
scabrous or with 0.3-0.8 mm hairs; lemmas 5.5-7.5 mm, lanceolate,
glabrous or with 0.3-0.8 mm hairs, sharply acute or awned, awns 0.8-3.5
mm; paleas 5.8-7.6 mm, scabrous, acute; anthers 2.5-5.1 mm; lodicules 1.3-1.5
mm. Caryopses 4.3-5 mm. 2n = 14, rarely 28.
Psathyrostachys juncea is native to central Asia, primarily to the
Russian and Mongolian steppes. It was introduced into North America as
a forage grass. It has become established at various locations from the
Yukon Territory through the prairie provinces to Arizona. It is drought
resistant and tolerant of saline soils. In its native range, it grows on
stony slopes and adjacent to roads,at elevations up to 5500 m.
Psathyrostachys juncea closely resembles Leymus cinereus, differing primarily in having a rachis that breaks up at maturity. Immature plants can be identified by their shorter ligules and the more uniform appearance of the spikelets compared to Leymus cinereus. Plants with pilose florets have been treated as a distinct taxon; such recognition is not merited.