| Gerrit Davidse |
Plants annual or perennial;
cespitose, sometimes stoloniferous, occasionally with rhizomes. Culms 2-60
cm. Sheaths
open, throats glabrous or long-ciliate; ligules of hairs, or membranous
and ciliate, or ciliolate; blades flat or rolled, glabrous or villous. Inflorescences
terminal, sometimes also axillary, panicles, spikes, or racemes. Spikelets
with 2-5(10) florets, distal florets reduced; disarticulation above
the glumes and between the florets. Glumes exceeding or exceeded
by the florets, herbaceous, 3-5-veined, glabrous or scabridulous, sometimes
with stiff, papillose-based hairs, glume apices acute or acuminate; calluses glabrous; lemmas
herbaceous, with acute or clavate hairs in marginal rows or variously scattered,
lemma apices unlobed, acute to long-acuminate; paleas sometimes
with tufts of hairs on the margins; lodicules 2, obtriangular,
glabrous or with bristles, 1-3-veined, lodicule apices at least as thick
as the base; anthers 3, 0.3-3.1
mm. Caryopses 0.7-2.2 mm long, 0.4-1.1 mm wide; pericarp poorly
separable, dull, smooth or rugulose, glabrous; embryos 1/3-1/2
as long as the caryopses; hila punctiform. x = 6. Desvaux
did not explain the etymology of Tribolium. It may be derived
from the Greek trilobos,
a name of various prickly plants (Quattrocchi 1999) or the Latin tria,
three, and bolus, a fiery meteor in the form of an arrow, referring
to the appearance of the mature spikelets with their three florets and
coarsely hairy glumes, which, like the lemmas, may be awned or acuminate
(Clifford 1996).
Tribolium is a southern African genus of 10 species. It is unusual in the
tribe in having unlobed lemmas, but has the haustorial synergids, bilobed or bi-awned
prophylls, and stalked ovaries characteristic of the tribe. Linder and Davidse
(1997) suggested that its closest relatives are probably Schismus,
Karroochloa, and Rytidosperma.
1. Tribolium obliterum (Hemsley) Renvoize
Plants perennial; usually cespitose, sometimes stoloniferous, stolons to
30 cm. Culms 12-40 cm, decumbent or erect, branching intravaginal. Sheaths
mostly glabrous, ciliate distally; ligules about 0.2 mm, membranous, ciliolate;
blades 1-8 cm long, 0.5-1 mm wide. Inflorescences terminal, panicles,
1-5 cm long, 5-15 mm wide, obovate or cylindrical, with 10-40 spikelets; branches
scabridulous. Spikelets 3.5-4.5 mm, with 5-10 florets. Glumes 2.5-3.5
mm, longer than the basal lemmas but exceeded by the distal florets, 5-veined,
glabrous or scabridulous over the veins, acute; lemmas 2-3.5 mm, with submarginal
tufts of hairs in the lower 1/2, hairs acute, 0.2-0.3 mm, lemma apices acute to
acuminate; paleas 1.5-2 mm, glabrous between the keels, sometimes with
tufts of hairs on the margins; anthers 0.3-0.5 mm, pale yellow. Caryopses
obovate, 0.7-1 mm long, 0.5-0.7 mm wide, plano-convex to concavo-convex, pale
brown, smooth; embryos 0.4-0.5 mm; hila 0.1 mm. 2n = 24,
36.
Tribolium obliterum is native to Cape Province, South Africa, where it
usually grows in gravelly, well-drained soils at elevations below 600 m. It has
been introduced into Australia and St. Helena and was recently discovered in a
roadside ditch near Fort Ord, Monterey County, California. It appears to be naturalized
there.