17.39   WILLKOMMIA Hack.
J.K. Wipff

Plants annual or perennial; cespitose or tufted, often stoloniferous. Culms 12-80 cm, herbaceous, unbranched distally. Leaves involute to flat; ligules membranous, ciliate. Inflorescences terminal, panicles of several, racemosely arranged, spikelike branches, mostly exceeding the leaves, lower branches sometimes partially included in the upper leaf sheaths at maturity; branches not woody, unilateral, with 2 rows of appressed, imbricate, solitary spikelets, terminating in a rudimentary, sterile spikelet, sterile spikelets sometimes consisting of 1 or 2 small scales. Spikelets dorsally compressed, with 1 floret; disarticulation below the glumes. Lower glumes to 2/3 as long as the spikelets, veinless; upper glumes equaling the florets, 1-veined; calluses acute to pointed; lemmas thinly membranous, 3-veined, apices rounded to acute, mucronate, shortly awned, or unawned. Caryopses ellipsoid. x = 10. Named for Heinrich Moriz Willkomm (1821-1895), a German botanist and explorer.

Willkommia is a genus of four species, three native to southern tropical Africa and one to the Americas, including the Flora region.


1.   Willkommia texana Hitchc.
Willkommia

Plants perennial; cespitose or stoloniferous. Culms (12)17-65 cm, glabrous; nodes 4-6(7), glabrous. Leaves cauline; sheaths mostly glabrous, upper margins sometimes pubescent; ligules 0.3-0.6 mm, truncate; blades (1)4-21 cm long, 0.6-3.5 mm wide, involute to flat, glabrous, margins pubescent. Panicles (6)7-34(44) cm long, 4-10 mm wide; branches 4-13(20), 1-8 cm, appressed, each branch usually terminating in a reduced, sterile spikelet, lowest branches usually partially enclosed in the uppermost sheaths at maturity. Spikelets 3.5-5 mm; disarticulation usually below the glumes, sometimes the first glume persistent. Lower glumes 2-3.5 mm, glabrous; upper glumes 3.3-4.8 mm, glabrous and smooth below, scabridulous above; lemmas 3.0-4.2 mm, usually appressed pilose between the veins, lateral veins obscure, midveins sometimes excurrent; paleas 2.5-3.4 mm, pubescent between the veins; anthers 3, 0.3-0.9(1.1) mm, yellow-orange. Caryopses 1.4-2 mm. 2n = 60.

Willkommia texana grows on clay pans, alkaline flats, and sandy soils, in open or bare areas of Texas and (rarely) Oklahoma and, as a disjunct, in Argentina. The Oklahoma record may represent a recent introduction. North American plants belong to Willkommia texana Hitchc. var. texana. They differ from the Argentinean variety, W. texana var. stolonifera Parodi, in being cespitose rather than stoloniferous.