| J.K. Wipff |
Plants annual or perennial; cespitose
or tufted, often stoloniferous. Culms 12-80 cm, herbaceous, unbranched
distally. Leaves involute to flat; ligules membranous, ciliate.
Inflorescences terminal, panicles of several, racemosely arranged, spikelike
branches, mostly exceeding the leaves, lower branches sometimes partially included
in the upper leaf sheaths at maturity; branches not woody, unilateral,
with 2 rows of appressed, imbricate, solitary spikelets, terminating in a rudimentary,
sterile spikelet, sterile spikelets sometimes consisting of 1 or 2 small scales.
Spikelets dorsally compressed, with 1 floret; disarticulation below
the glumes. Lower glumes to 2/3 as long as the spikelets, veinless; upper
glumes equaling the florets, 1-veined; calluses acute to pointed; lemmas
thinly membranous, 3-veined, apices rounded to acute, mucronate, shortly awned,
or unawned. Caryopses ellipsoid. x = 10. Named for Heinrich Moriz
Willkomm (1821-1895), a German botanist and explorer.
Willkommia is a genus of four species, three native to southern tropical
Africa and one to the Americas, including the Flora region.
1. Willkommia texana Hitchc.
Willkommia
Plants perennial; cespitose or stoloniferous. Culms (12)17-65 cm,
glabrous; nodes 4-6(7), glabrous. Leaves cauline; sheaths
mostly glabrous, upper margins sometimes pubescent; ligules 0.3-0.6 mm,
truncate; blades (1)4-21 cm long, 0.6-3.5 mm wide, involute to flat, glabrous,
margins pubescent. Panicles (6)7-34(44) cm long, 4-10 mm wide; branches
4-13(20), 1-8 cm, appressed, each branch usually terminating in a reduced, sterile
spikelet, lowest branches usually partially enclosed in the uppermost sheaths
at maturity. Spikelets 3.5-5 mm; disarticulation usually below the
glumes, sometimes the first glume persistent. Lower glumes 2-3.5 mm, glabrous;
upper glumes 3.3-4.8 mm, glabrous and smooth below, scabridulous above;
lemmas 3.0-4.2 mm, usually appressed pilose between the veins, lateral
veins obscure, midveins sometimes excurrent; paleas 2.5-3.4 mm, pubescent
between the veins; anthers 3, 0.3-0.9(1.1) mm, yellow-orange. Caryopses
1.4-2 mm. 2n = 60.
Willkommia texana grows on clay pans, alkaline flats, and sandy soils,
in open or bare areas of Texas and (rarely) Oklahoma and, as a disjunct, in
Argentina. The Oklahoma record may represent a recent introduction. North American
plants belong to Willkommia texana Hitchc. var. texana.
They differ from the Argentinean variety, W. texana var. stolonifera
Parodi, in being cespitose rather than stoloniferous.